Staff+ Application Security Engineer at Anthropic

Hybrid - Remote-Friendly (Travel-Required), San Francisco, CA, Seattle, WA, New York City, NY

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This senior Application Security Engineer role at Anthropic focuses on securing the AI systems that build, serve, and extend Claude. Leveraging Claude as the primary tool, the engineer will design and build LLM-powered security systems, lead threat modeling and secure design reviews, evolve the bug bounty program, collaborate with cross‑functional teams, and participate in on‑call rotations for incident response and launch consults.

Salary

USD 320,000 - 485,000

Requirements

Skills

  • Hands-on application and infrastructure security experience, including cloud and containerized environments
  • Production-quality coding ability in at least one of Python, Go, Rust, or TypeScript
  • Practical threat-modeling and vulnerability-identification skills
  • Demonstrated ability to operate with high autonomy and ambiguity
  • Clear technical communication with both engineers and leadership

Responsibilities

  • Design, build, and operate Claude-powered security systems — LLM-driven code analysis, automated vulnerability remediation, AI-assisted threat modeling — and own one or more of them end-to-end
  • Lead secure design reviews and threat modeling for novel AI systems, identifying risks that don't map to existing frameworks
  • Evolve a public bug bounty program where automation handles routine triage and root-cause work, and engineers handle escalations and corner cases
  • Partner with Product, Infrastructure, and Research teams as an embedded security owner — consulting on launches, shaping architecture, and influencing decisions where security is the constraint
  • Share an operational on-run rotation with the rest of the team — bounty escalations, incident response, and launch consults on systems serving Claude in production

Technologies

Claude (LLM)PythonGoRustTypeScriptCloud environmentsContainerizationStatic analysisAI-assisted threat modelingBug bounty automationSecurity automationSandboxing

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